New York Natural Heritage Program
Nine-spotted Lady Beetle
Coccinella novemnotata Herbst, 1793

Arnett, Jr., Ross H., ed. 1983. Checklist of the Beetles of North and Central America and the West Indies. Flora and Fauna Publications, Gainesville, Florida. 24 P. (Pertains to all subsequent fasicle updates as well).
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Harmon, J.P., E. Stephens, and J. Losey. 2007. The decline of native coccinellids (Coleoptera: Coccinellidae) in the United States and Canada. Journal of Insect Conservation. 11: 85-94.
Hessler, Louis S., Ginger McNickle, Michael A. Catangui, John E. Losey, Eric A. Beckendorf, Leonard Stellwag, Danielle M. Brandt, and Pamela B. Bartlett. 2012. Method for continuously rearing Coccinella lady beetles (Coleoptera: Coccinellidae). The Open Entomology Journal 6:42-48.
Ijaz, D. 2013. "Coccinella novemnotata" (On-line), Animal Diversity Web. Accessed December 29, 2013 at">
Keys, Jr.,J.; Carpenter, C.; Hooks, S.; Koenig, F.; McNab, W.H.; Russell, W.;Smith, M.L. 1995. Ecological units of the eastern United States - first approximation (cd-rom), Atlanta, GA: U.S. Department of Agriculture, Forest Service. GIS coverage in ARCINFO format, selected imagery, and map unit tables.
Losey, J., J. Perlman, J. Kopco, S. Ramsey, L. Hesler, E. Evans, L. Allee, R. Smyth. 2012. Potential causes and consequences of decreased body size in field populations of">Coccinella novemnotata. Biological Control, 61: 98-103.
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NatureServe. 2015. NatureServe Explorer: An online encyclopedia of life [web application]. Version 7.1. NatureServe, Arlington, Virginia. Available 
Stephens, Erin and John Losey. 2003. The decline of C-9- New York State’s insect. The Xerces Society. Wings: Essays on Invertebrate Conservation. Fall 2003 pp. 8-12.
The Xerces Society. 2013. “Ladybird beetles: nine-spotted lady beetle (Coccinella novemnotata).” the Xerces Society for Invertebrate Conservation."> (date accessed: December 29, 2013).


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This guide was authored by: Hollie Y. Shaw
Information for this guide was last updated on: 07-Aug-2017