New York Natural Heritage Program
Nine-spotted Lady Beetle
Coccinella novemnotata Herbst, 1793
Insects

References
Arnett, Jr., Ross H., ed. 1983. Checklist of the Beetles of North and Central America and the West Indies. Flora and Fauna Publications, Gainesville, Florida. 24 P. (Pertains to all subsequent fasicle updates as well).
Cornell University. 2013. “The Lost Ladybug Project.” http://www.lostladybug.org/">www.lostladybug.org. (date accessed: December 29, 2013).
Gordon, R. 1985. The http://animaldiversity.ummz.umich.edu/accounts/Coccinellidae/">Coccinellidae (http://animaldiversity.ummz.umich.edu/accounts/Coleoptera/">Coleoptera) of America north of Mexico. Journal of the New York Entomological Society, 93: 1-912. 
Harmon, J.P., E. Stephens, and J. Losey. 2007. The decline of native coccinellids (Coleoptera: Coccinellidae) in the United States and Canada. Journal of Insect Conservation. 11: 85-94.
Hessler, Louis S., Ginger McNickle, Michael A. Catangui, John E. Losey, Eric A. Beckendorf, Leonard Stellwag, Danielle M. Brandt, and Pamela B. Bartlett. 2012. Method for continuously rearing Coccinella lady beetles (Coleoptera: Coccinellidae). The Open Entomology Journal 6:42-48.
Ijaz, D. 2013. "Coccinella novemnotata" (On-line), Animal Diversity Web. Accessed December 29, 2013 at http://animaldiversity.ummz.umich.edu/accounts/Coccinella_novemnotata/">http://animaldiversity.ummz.umich.edu/accounts/Coccinella_novemnotata/
Keys, Jr.,J.; Carpenter, C.; Hooks, S.; Koenig, F.; McNab, W.H.; Russell, W.;Smith, M.L. 1995. Ecological units of the eastern United States - first approximation (cd-rom), Atlanta, GA: U.S. Department of Agriculture, Forest Service. GIS coverage in ARCINFO format, selected imagery, and map unit tables.
Losey, J., J. Perlman, J. Kopco, S. Ramsey, L. Hesler, E. Evans, L. Allee, R. Smyth. 2012. Potential causes and consequences of decreased body size in field populations of http://animaldiversity.ummz.umich.edu/accounts/Coccinella_novemnotata/">Coccinella novemnotata. Biological Control, 61: 98-103.
NatureServe. 2005. NatureServe Central Databases. Arlington, Virginia. USA
NatureServe. 2015. NatureServe Explorer: An online encyclopedia of life [web application]. Version 7.1. NatureServe, Arlington, Virginia. Available http://www.natureserve.org/explorer. 
Stephens, Erin and John Losey. 2003. The decline of C-9- New York State’s insect. The Xerces Society. Wings: Essays on Invertebrate Conservation. Fall 2003 pp. 8-12.
The Xerces Society. 2013. “Ladybird beetles: nine-spotted lady beetle (Coccinella novemnotata).” the Xerces Society for Invertebrate Conservation. http://www.xerces.org/nin-spot-ladybird-beetle/">http://www.xerces.org/nin-spot-ladybird-beetle/ (date accessed: December 29, 2013).

Acknowledgements

Funding from the NYSDEC Hudson River Estuary Program made the initial development of these guides possible.

Additional support has been provided by

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Modifications and updates: David Marston and Matthew Buff.

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This guide was authored by: Hollie Y. Shaw
Information for this guide was last updated on: 07-Aug-2017