New York Natural Heritage Program
Eastern Small-footed Myotis
Myotis leibii (Audubon and Bachman, 1842)

Baker, R. H. 1983. Michigan mammals. Michigan State University Press. 642 pp.
Banfield, A. W. F. 1974. The mammals of Canada. University of Toronto Press, Toronto, Canada. 438 pp.
Barbour, R. W., and W. H. Davis. 1969. Bats of America. The University of Kentucky Press, Lexington, Kentucky.
Best, T. L., and J. B. Jennings. 1997. Myotis leibii. Mammalian Species (547):1-6.
Caire, W., J. D. Tyler, B. P. Glass, and M. A. Mares. 1989. Mammals of Oklahoma. University of Oklahoma Press, Norman. Oklahoma. 567 pp.
Chambers, R.E. 1983. Integrating timber and wildlife management. State University of New York, College of Environmental Science and Forestry and New York State Department of Environmental Conservation.
Dalton, Virginia M. 1987. Distribution, Abundance, and Status of Bats Hibernating in Caves in Virginia. Virginia Journal of Science 38(4): 369-379.
Fenton, M. B. 1972. Distribution and over-wintering of Myotis leibii and Eptesicus fuscus (Chiroptera: Vespertilionidae) in Ontario. Royal Ontario Museum,, Life Sciences Occas. Pap. (21):1-8.
Godin, A. J. 1977. Wild mammals of New England. Johns Hopkins University Press, Baltimore. 304 pp.
Gordon, D.C. 1986. Mammals of Jefferson and Lewis counties, New York. Humphrey Press, Canandaigua, N.Y. 135 pp.
Hall, E. Raymond. 1981. The Mammals of North America, Vols. I & II. John Wiley & Sons, New York, New York. 1181 pp.
Hamilton, W.J., Jr. and J.O. Whitaker, Jr. 1979. Mammals of the eastern United States. Cornell University Press. Ithaca, New York. 346 pp.
Handley, C. O., Jr. 1991. Mammals. Pages 539-616 in K. Terwilliger, coordinator. Virginia's endangered species: proceedings of a symposium. McDonald and Woodward Publishing Company, Blacksburg, Virginia.
Harvey, M.J., J.S. Altenbach, and T.L. Best. 1999. Bats of the United States. Arkansas Game and Fish Commission and United States Fish and Wildlife Service, Little Rock, Arkansas, 64 pp.
Herd, R. M. 1987. Electrophoretic divergence of Myotis leibii and Myotis ciliolabrum (Chiroptera: Vespertilionidae). Can. J. Zool. 65:1857-1860.
Hitchcock, H. B. 1955. A summer colony of the least bat Myotis subulatus leibii (Audubon and Bachman). Canadian Field-Naturalist 69(2):31.
Hitchcock, H. B. 1965. Twenty-three years of bat banding in Ontario and Quebec. Canadian Field Naturalist 79(1):4-14.
Johnson, Joshua B., and J. Edward Gates. 2007. Food habits of Myotis leibii during fall swarming in West Virginia. Northeastern Naturalist 14: 317-322.
Jones, J. K., Jr., R. S. Hoffman, D. W. Rice, C. Jones, R. J. Baker, and M. D. Engstrom. 1992a. Revised checklist of North American mammals north of Mexico, 1991. Occasional Papers, The Museum, Texas Tech University, 146:1-23.
Keys, Jr.,J.; Carpenter, C.; Hooks, S.; Koenig, F.; McNab, W.H.; Russell, W.;Smith, M.L. 1995. Ecological units of the eastern United States - first approximation (cd-rom), Atlanta, GA: U.S. Department of Agriculture, Forest Service. GIS coverage in ARCINFO format, selected imagery, and map unit tables.
Menzel, J. M., M. A. Menzel, W. M. Ford, J. W. Edwards, S. R. Sheffield, J. C. Kilgo, and M. S. Bunch. 2003. The distribution of the bats of South Carolina. Southeastern Naturalist 2:121-152.
Merritt, J. F. 1987. Guide to the mammals of Pennsylvania. University of Pittsburgh, Pittsburgh, PA.
Moosman Jr, Paul R., Howard H. Thomas, and Jacques Pierre Veilleux. 2007. Food habits of eastern small-footed bats (Myotis leibii) in New Hampshire.The American Midland Naturalist 158: 354-360.
NYSDEC [online]. 2012. DEC Reports: 2012 Winter Bat Survey Results. Department of Environmental Conservation. <> (3 February 2014).
NatureServe. 2005. NatureServe Central Databases. Arlington, Virginia. USA
New York State Department of Environmental Conservation, Division of Fish, Wildlife, and Marine Resources. 2006. New York State Comprehensive Wildlife Conservation Strategy. Albany, NY: New York State Department of Environmental Conservation.
New York State Department of Environmental Conservation. Checklist of the amphibians, reptiles, birds, and mammals of New York State, including their protective status. Nongame Unit, Wildlife Resources Center, Delmar, NY.
Schwartz, C. W., and E. R. Schwartz. 1981. The wild mammals of Missouri. University of Missouri Press, Columbia. 356 pp.
Tuttle, M. D. 1964. Myotis subulatus in Tennessee. J. Mamm. 45:148-149.
Tuttle, M.O. 1974. An improved trap for bats. J. Mammal. 55(2):475-477.
U.S. Fish and Wildlife Service. 2011. 90-Day finding on a petition to list the eastern small-footed bat and the northern long-eared bat as threatened or endangered. Vol. 76 No. 125, Department of the Interior.
U.S. Fish and Wildlife Service. 2013. 12-Month finding on a petition to list the eastern small-footed bat and the northern long-eared bat as threatened or endangered; Listing the northern long-eared bat as an endangered species; Proposed rule. Vol. 78 No. 191, Department of the Interior.
Wilson, D. E., and D. M. Reeder (editors). 1993. Mammal Species of the World: a Taxonomic and Geographic Reference. Second Edition. Smithsonian Institution Press, Washington, DC. xviii + 1206 pp. Available online at:
Wimsatt, W. A. 1945. Notes on breeding behavior, pregnancy, and parturition in some vespertilionid bats of the eastern United States. Jounal of Mammalogy 26(1):23-33.
van Zyll de Jong, C. G. 1984. Taxonomic relationships of Nearctic small-footed bats of the Myotis leibii group (Chiroptera: Vespertilionidae). Can. J. Zool. 62:2519-2526.
van Zyll de Jong, C. G. 1985. Handbook of Canadian Mammals. Volume 2. Bats. National Museums of Canada, Ottawa, Ontario, Canada. 212 pp.


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Information for this guide was last updated on: 08-Aug-2017