A plant of two different areas and habitats in New York. In the Hudson Valley, it is restricted to the edges of freshwater intertidal mudflats, sandy or rocky shorelines of tidal creeks and other waterways, edges of freshwater tidal marshes, and gravel shores along the freshwater tidal portions of the Hudson River. In western New York, it is found along gravelly streams through red maple-hardwood swamps of the Tonawanda Oak Orchard Swamp drainage (New York Natural Heritage Program 2004). Semi-aquatic, in marshes and along streams, especially on calcareous substrate (Gleason & Cronquist 1991). In and along streams and swampy woods (Fernald 1970).

Freshwater intertidal mudflats A sparsely vegetated community characterized by low rosette-leaved aquatics. This community occurs on exposed intertidal mudflats where the water is fresh (salinity less than 0.5 ppt). This community is best developed where mudflats are nearly level so that broad expanses are exposed at low tide. The plants are completely submerged in 0.9 to 1.2 m (3 to 4 ft) of water at high tide and they are usually coated with mud. Freshwater intertidal shore A community of the intertidal gravelly or rocky shores of freshwater tidal rivers and creeks, sometimes occurring at the base of cliffs. The vegetation may be very sparse. Freshwater tidal creek The aquatic community of a shallow, tidally flooded freshwater creek with submerged areas averaging less than 2 m (6 ft) deep at low tide.

Freshwater tidal marsh A marsh community that occurs in shallow bays, shoals, and at the mouth of tributaries of large tidal river systems, where the water is usually fresh (salinity less than 0.5 ppt), and less than 2 m (6 ft) deep at high tide. Typically there are two zones in a freshwater tidal marsh: a low-elevation area dominated by short, broadleaf emergents bordering mudflats or open water, and a slightly higher-elevation area dominated by tall grass-like plants. Marsh headwater stream The aquatic community of a small, marshy perennial brook with a very low gradient, slow flow rate, and cool to warm water that flows through a marsh, fen, or swamp where a stream system originates. These streams usually have clearly distinguished meanders (i.e., high sinuosity) and are in unconfined landscapes. Red maple-hardwood swamp* A hardwood swamp that occurs in poorly drained depressions, usually on inorganic soils. Red maple is usually the most abundant canopy tree, but it can also be codominant with white, green, or black ash; white or slippery elm; yellow birch; and swamp white oak. * probable association but not confirmed

Red Maple (Acer rubrum var. rubrum) Silver Maple (Acer saccharinum) Broad-leaved Water-plantain (Alisma subcordatum) Waterhemp Pigweed (Amaranthus cannabinus) False Indigobush (Amorpha fruticosa) American Hogpeanut (Amphicarpaea bracteata) Maryland Bur-marigold (Bidens bidentoides) Nodding Beggar-ticks (Bidens cernua) Broad Waterweed (Elodea canadensis) Common Sneezeweed (Helenium autumnale) Yellowseed False Pimpernel (Lindernia dubia) Cardinal Flower (Lobelia cardinalis) Marsh Seedbox (Ludwigia palustris)

Fringed Loosestrife (Lysimachia ciliata) Golden Club (Orontium aquaticum) Green Arrow-arum (Peltandra virginica) Springs Clearweed (Pilea fontana) Canada Clearweed (Pilea pumila) Pickerelweed (Pontederia cordata) Native Blackeyed Susan (Rudbeckia hirta) Grassleaf Arrowhead (Sagittaria graminea) Broadleaf Arrowhead (Sagittaria latifolia) Water Chestnut (Trapa natans) Riverbank Grape (Vitis riparia) Eastern Wild Rice (Zizania aquatica)